In 1885, Shostka, Russia, the Mousekewitzes-- a Russian-Jewish family members of computer mice-- who deal with a human family named Moskowitz are having an event of Hanukkah where Papa offers his hat to his child, Fievel, and tells of a fantastic location called America, where there are no felines. The party is disturbed when a battery of Cossacks ride through the town square in an arson strike as well as their pet cats furthermore assault the village computer mice.
In Hamburg, Germany, the Mousekewitzes board a vagrant cleaner went to America. The crossing shows itself long and also burdensome, along with the vagrant cleaner being buffetted by an angry sea. Throughout a thunderstorm, Fievel all of a sudden finds himself divided from his family members as well as washed too far. Believing that he has sunk, they proceed to New York City as planned, though they come to be depressed at his loss.
Nevertheless, Fievel drifts to The U.S.A. in a bottle and also, after a pep talk from a French pigeon called Henri, starts a quest to discover his family. He is waylaid by conman Warren T. Rat, that acquires his trust and then sells him to a sweatshop. He gets away with Tony Toponi, a street-smart Italian mouse, and they join up with Bridget, an Irish computer mouse attempting to rouse her fellow mice to take on the pet cats. When a gang of them called the Mott Road Maulers strikes a mouse market, the immigrant computer mice find out that the tales of a cat-free nation are not real.
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